Phegea

driemaandelijks tijdschrift van de

VLAAMSE VERENIGING VOOR ENTOMOLOGIE

Afgiftekantoor 2 1 70 Merksem 1 ISSN 077 1 -5277

Periode: april - mei “juni 2009 Erkenningsnr. P209674

Redactie: Dr. J.-P. Borie (Compiègne, France), Dr. L. De Bruyn (Antwerpen), T. C. Garrevoet
(Antwerpen), B. Goater (Chandlers Ford, England), Dr. K. Maes (Tervuren), Dr. K. Martens
(Brussel), H. van Oorschot (Amsterdam), W. O. De Prins (Eeefdaal).

Redactie-adres: W. O. De Prins, Eeefdaal 40 1B, B-3061 Leefdaal (Belgium).
e-mail: willy.de.prins@telenet.be.

Jaargang 37, nummer 2 1 juni 2009

A new Feaella species (Pseudoscorpiones: Feaellidae)
from Kenya

Hans Henderickx1,2

Abstract. Feaella jocquei sp. n., a new pseudoscorpion from Kenya is described.
Samenvatting. Feaella jocquei sp. n., een nieuwe pseudoschorpioen uit Kenya wordt
beschreven.

Résumé. Feaella jocquei sp. n., une espèce nouvelle, provenante du Kénya, est décrite.

Key words. Pseudoscorpiones - Feaella jocquei - Tanzania - Kenya.

Henderickx, H.: 'Department of Biology, Universiteit Antwerpen (UA), Groenenborgerlaan 171,
2020 Antwerpen, Belgium. 2Royal Belgian Institute of Natural Sciences, Department
Entomology, Vautierstraat 29, B-1000 Brussels, (Adress for correspondence: Hemelrijkstraat
4, B-2400 Mol; cavexplorer@gmail.com).

Introduction

Feaella belongs to the Feaelloidea, which is, based on molecular data,
considered to represent the most basal pseudoscorpion clade (Murienne et al.
2008). Some Feaella occupy an isolated ecological pocket, f.i. Feaella leleupi
Beier, 1959 under stones on Ile Mboko, Lake Tanganyika (Leleup, personal
communication). Several other Feaella sp. have been found on their locus
typicus only and seem to be restricted to that area.

The present publication is about a Feaella species that is sieved from litter
near the Taita Discovery Center in Tsavo, Kenya. The species bears an anterior
row of 6 and a posterior row of 4 rounded protuberances on the carapace
(cucullus) and can be placed in the subgenus Feaella L(lingsen,; 1906. It is
morphologically and geographically close to Feaella ( Feaella)' mombasica
Beier, 1955 but it shows significant morphological differences with the latter and
all other Feaella species and is therefore described here as a new species.

Phegea 37 (2) (01.VI.2009)S41

Material and methods

A male and tritonymph (Fig. 3) of this species were collected by Jocqué and
Warui (Royal Museum for Central Africa, Tervuren) by sieving litter, then
transferred to ethanol 70%. Examination and measurements have been carried
out with a Leitz microscope, electron microscopy with the FEI Quanta-200.
Special attention was given to non-destructive examination techniques, no parts
were removed from the holotype and the ESEM scanning microscopy was
performed in low pressure water vapour.

All measurements are in mm; (length=L x width=W), the ratio is the
length/width index of an article.

Systematics

Feaella ( Feaella ) jocquei sp. n.

(Figs. 1, 2)

Type material: Male holotype and 1 tritonymph paratype, sample 209.608,
Royal Museum for Central Africa, Tervuren. Information on label: Kenya
28. III. 2000, loc. Tsavo, Taita Discovery Center, Eco. Acacia-Commiphora
forest, Maungu near entrance gate, sieving litter, ree. Jocqué R. & Warui, C.,
R.G. Mus. Afr. Centr. 209.608.

Etymology: Patronym in honour of Rudy Jocqué, who collected the
specimens.

Diagnosis: A typical Feaella {Feaella) species with four equally sized, well
developped eyes, the anterior part of the rear eye pair covered by carapacal
cuticula (Fig. 2b). Second tergal plate completely split, pedipalpal trochanter
with dorsal tubular projection. The new species is most closely related to Feaella
{Feaella) mombasica but differs from it by the split second tergal plate and the
four equally sized eyes.

Male holotype, description: (measurements in mm, ratio is L/W): Carapace
longer than wide (1.42 x), carapace and opisthosoma with a coarse reticulated
texture forming ridges (fig. 2b). Both pairs of eyes well developped, equal in
size. The anterior part of the second pair covered by carapace cuticula. Chelicera
large, galea simple without rami, pointed. Movable finger with round terminal
dimple and one seta, hand with 4 macrosetae. 6 microsetae near the edge of the
dorsal, reticulated zone of the cheliceral hand. The proximal and ventral part of
the chelicera lacks reticulation, that part is smooth as a hinge joint. Serrula of
movable finger with 17 lobes, First lobe pointed.

Tergal setae reduced, cannot be seen between the course reticulation. First
tergite narrow, adapted to the flexible carapace-opisthosoma connection that is
typical for Feaella. Tergites II to X distinct and completely split, XI th tergite
ventrally joined with the terminal stemite, the circular anus is situated in this
sclerite complex.

Phegea 37 (2) (01.VI.2009): 42

First and second stemites fused, third sternite a narrow undivided strip
bording the genital opening, extending laterally under the fourth coxa. Sternites
IV to X completely divided. Pedipalpal coxa (Fig. 2d) with lateral thom, near the
joint with the leg coxa narrowed and in this zone without coarse granulation.
The smooth surface there shows only a flat or honeycomb pattem, that might
facilitate movement of the special joint connection. A honeycomb pattem is also
visible on the part of the leg trochanter that moves in the leg coxa. (Fig. 2e).
Pleurites with two rows of 15 plates.

Pedipalp shape (Fig. la) typical Feaella, with a coarse reticulated structure.
This reticulation lacks on the terminal part of the fingers and in the zone where
the trichobothria are. Trochanter as long as broad, dorsally with a blunt 0.1 mm
x 0.03 mm tubular projection. Femur 1.75x as long as broad, with a ridge that
ends in a mediobasal thom. Tibia 3x as long as broad, widening distally. Hand
(without fingers) short, only slightly longer than broad (1.06x), complete chela
(Fig. lb) with pedicel 3,74x longer than broad. Both fingers intemal with a large
basal tooth (thom). Several longitudinal rows of unequal teeth ending in a
terminal teeth ‘crown’ of 5 teeth in both fingers. Fixed fmger with an extemal
row of 1 1 teeth, movable fmger with a ventral row of 1 5 teeth.

Fixed fmger with 8, movable with 4 trichobothriae, positions illustrated on
Fig. lb. The intemal trichobothriae on this extemal view are outlined with dots.

Legs monotarate, a movable joint between telofemur and basifemur.
Aroleum not longer than claws.

Measurements (mm): Body length with chelicera 2.36, without chelicera
2.20. Pedipalp: trochanter 0.16/0.16; femur 0.59/0.34; tibia 0.51/0.17; chela
(with pedicel) 0.59/0.16; hand 0.17/0.16; movable fmger L=0.42. Carapace
0.71/0.45. Anterior and posterior eyes equal in size (diameter is 0.067).

Leg 1 (Fig. Ie): trochanter 0.19/0.14; basifemur 0.24/0.08; telofemur
0.28/0.10; tibia 0.21/0.08; tarsus 0.38/0.04.

Leg IV (Fig. ld): trochanter 0.37/0.17; basifemur 0.23/0.12; telofemur
0.45/0.16; tibia 0.53/0.09; tarsus 0.57/0.05.

Discussion

Eight species and one subspecies of Feaella are described from Africa. On
base of the frontal carapacal lobes a further separation in three subgenera has
been made: Feaella (Ellingsen , 1906), Tetrafeaella (Beier, 1955) and Difeaella.
(Beier, 1966). Feaella jocquei n. sp. belongs to the subgenus Feaella (frontal
edge of the carapace with 6 lobes) but differs clearly from the other species of
this subgenus. The fixed fmger bears no pronounced dorso-lateral projection as
in Feaella {Feaella) mirabilis Ellingsen, 1906, the new species can also be
separated from F. {F.) mirabilis by the long tubular structure on the dorsal side
of the trochanter (Fig. 2c).

Phegea 37 (2) (01.VI.2009): 43

Fig. 1. Feaella jocquei sp.n., holotypus: a - habitus; b - chela, antiaxal lateral view; c- leg I; d. leg
IV.

Phegea 37 (2) (01.VI.2009): 44

Fig. 2. Feaella jocquei sp.n., holotypus, ESEM microscopy: a. right celicera; b- carapace, First and
second tergite; c - trochanter, lateral; d- coxa of pedipalp; e - coxa of legs.

Phegea 37 (2) (01.VI.2009): 45

Fig. 3. Feaella jocquei sp. n., tritonymph, ESEM microscopy: a. carapace and First tergites; b-
chela, antiaxal lateral view; c - right pedipalp, dorsal view

Phegea 37 (2) (01.VI.2009): 46

Feaella ( Feaella ) mombasica has such a tubular structure on the trochanter,
but unlike F. mombasica the new species has a completely split second tergite
pair. The frontal eyes of F. mombasica are significant smaller compared to the
second pair while both pairs of eyes of F. jocquei sp. n. are the same size. F.
jocquei sp. n. is larger than F. mombasica and the chelae are less slender. The
tritonymph in sample 209.608 from the same location and date has analogue
characteristics and will be subject to future examination.

Biology

F. jocquei sp. n. was sieved from litter in an Acacia-Commiphora forest, F.
mirabilis is exclusively corticole (under bark) (Heurtault-Rossi & Jézéquel
1965) and F. mombasica occurs in litter under bushes near the sea (Beier 1955).
The ability to occupy different ecological niches can contribute to the formation
of new species.

Distribution

Found on the type locality (Tsavo, Kenya) only. The presence of a new
Feaella species in Tsavo that is taxonomically and geographically close (190
km) to Feaella mombasica from Bamburi Beach, Mombasa, confirms that
Feaella populations in Africa can present high endemism.

Acknowledgements

The author is grateful to Dr Rudy Jocque (Royal Museum for Central Africa)
who offered valuable help and access to the arachnid collections, to Dr Herman
Goethals and Julien Cillis (Royal Belgian Institute of Natural Sciences, Brussel)
for support with the FEI Quanta-200 electron microscope.

References

Beier, M. 1955. Pseudoscorpionidea, gesammelt wahrend der schwedischen Expeditionen nach
Ostafrika 1937-38 und 1948. — Arkiv för Zoologi (2)7: 527-558.

Beier, M. 1966. Erganzungen zur Pseudoscorpioniden-Fauna des südlichen Afrika. — Annals of the
Natal Museum 18: 455-470.

Ellingsen, E. 1906. Report on the pseudoscorpions of the Guinea Coast (Africa) collected by
Leonardo Fea. — Annali del Museo Civico di Storia Naturale di Genova (3)2: 243-265.
Heurtault-Rossi, J. and Jézéquel, J. F. 1965. Observations sur Feaella mirabilis Eli. (Arachnide,
Pseudoscorpion). Les chélicères et les pattes-machoires des nymphes et des adultes. Description
de 1'appareil reproducteur. — Bulletin du Muséum National d'Histoire Naturelle , Paris (2) 37:
450-461.

Murienne, J., Harvey, M. S., Giribet, G. 2008. First molecular phylogeny of the major clades of
Pseudoscorpiones (Arthropoda: Chelicerata). — Molecular Phylogenetics and Evolution 49:
170-184.

Phegea 37 (2) (01.VI.2009): 47

Palloptera scutellata (Diptera: Pallopteridae) new for
the fauna of Belgium

John T. Smit, Jan Willem A. van Zuijlen & Bart Hanssens

Samenvatting. Palloptera scutellata (Diptera: Pallopteridae) nieuw voor de Belgische
fauna.

Een exemplaar van Palloptera scutellata (Macquart, 1835) werd gedetermineerd aan de hand van
een foto. Deze soort werd nog niet eerder gemeld uit België. Het totaal aantal uit België bekende
soorten Pallopteridae komt hiermee op acht.

Résumé. Palloptera scutellata (Diptera: Pallopteridae) nouvelle espèce pour la faune beige.
Un exemplaire de Palloptera scutellata (Macquart, 1835) a été déterminé a 1'aide d'une photo.
Cette espèce n'a jamais été mentionnée de Belgique. Le nombre d'espèces de Pallopteridae
connues de Belgique monte ainsi a huit.

Key words: Palloptera scutellata - faunistics - Belgium - First record.

Smit, J. T.: European Invertebrate Survey - the Netherlands, PO Box 9517, 2300 RA, Leiden,
the Netherlands. smitj@naturalis.nl

van Zuijlen, J. W. A.: Mendelssohnstraat 62, 5144 GH Waalwijk, the Netherlands.
jewe.vz@lycos.nl

Hanssens, B.: Natuurpunt Brussel en Milieu Commissie Brussel en Omgeving vzw
(www.cebe.be), Stroobantsstraat 35, 1 140 Brussel, Belgium. bhanssens@skynet.be

As in many other countries, the family Pallopteridae has received little
attention in Belgium. It is a family of medium-sized (2. 5-7.0 mm), greyish,
reddish to brownish yellow or pale yellow, rather sparsely bristled, acalyptrate
flies. This family can be distinguished by the following combination of
characters: head with one pair of reclinate orbital setae; vibrissae absent. Wings
maculated in most species, costa with subcostal break. Legs without dorsal
preapical setae on tibiae. Females with an elongate, piereer- type ovipositor,
varying from narrow to broad. Larvae have been found in sterns of Apiaceae,
Poaceae, Juncaceae, in flower heads of Asteraceae, or under bark of various
trees. The larvae living under bark are thought to be feeding on beetles or
mycelium. In general little is known about the biology of most species. The
family is represented by only a limited number of species in North-westem
Europe and up till now, only seven species were recorded from Belgium
(Gosseries 1991).

On February 10th 2008 a single female specimen of Palloptera scutellata
(Macquart, 1835) (fig. 1) was photographed in a garden in Brussels by the third
author. This is the first record of this species for Belgium. P. scutellata can be
easily distinguished from all other Palloptera species known from Belgium by
the dusted blueish-grey thorax, the orange-brown abdomen and the wing pattem
(fig. 2). This species has only recently been found in Germany (Stuke & Merz
2005) and the Netherlands (De Jong & Van Aartsen 2007).

Phegea 37 (2) (01.VI.2009): 48

Figure 1. Female Palloptera scutellata taken at a garden in Brussels. Photo B. Hanssens.

Figure 2. Female Palloptera scutellata taken at Doom, the Netherlands. The typical blueish-grey
thorax, orange-brown abdomen and wing pattem are clearly visible. Photo J. T. Smit.

Phegea 37 (2) (01.VI.2009): 49

The recent increase of records from the Netherlands in 2007 and 2008
suggests that this species has probably been present for a longer period and has
been largely overlooked due to its host plant and its aberrant flight period (Van
Eek et al. 2008). P. scutellata is associated with Juncus spp. and is mainly found
in stands of J. effusus. The species has been recorded in early spring, from
March until May/June (also one record from July), and in late autumn, from
October until December. It is suggested that the adults emerge in late autumn
and hibemate as adults, which would explain the aberrant flight period. The
large numbers observed in March in the Netherlands seem to confirm this
hypothesis. Another curious aspect of this species is the fact that nearly all
records are of females (Van Eek et al. 2008, De Jong & Van Aartsen 2007,
Stubbs 1969, Stuke & Merz 2005). This leads De Jong & Van Aartsen (2007) to
suspect that this species probably reproduces parthenogenetically. Whether this
is true or not remains unknown up till now and can only be refuted by finding
males.

References

Gosseries J. 1991. Pallopteridae. - In: Grootaert P., De Bruyn L. & M. De Meyer (eds.). Catalogue of
the Diptera of Belgium. — Studiedocumenten van het Koninklijk Belgisch Instituut voor
Natuurwetenschappen 70: 140.

De Jong H. & B. van Aartsen 2007. Palloptera scutellata (Diptera: Pallopteridae) in The
Netherlands. — Entomologische Berichten , Amsterdam 67: 105-107.

Stubbs A. E. 1969. Observations on Palloptera scutellata (Mcq) in Berkshire and Surray and
discussion on the larval habitats of British Pallopteridae (Diptera). — Entomologist’s Monthly
Magazine 104: 157-160.

Stuke J.-H. & B. Merz 2005. Drei fur Deutschland neu nachgewiesenen acalyptrate Fliegen (Diptera:
Lauxaniidae, Pallopteridae, Ulidiidae). — Studia dipterologica 12: 242-254.

Van Eek A., Zuijlen, J. W. A. van & J. T. Smit 2008. Palloptera scutellata (Macquart, 1835)
(Diptera, Pallopteridae) een recente aanwinst of lange tijd over het hoofd gezien. -
Vliegenmepper 17(1): 7-10.

Phegea 37 (2) (01.VI.2009): 50

Eupeodes goeldlini (Diptera: Syrphidae) new to the
Belgian fauna

Frank Van de Meutter

Abstract. Following a recent revision of Eupeodes and E. bucculatus (Rondani, 1857) in
particular, this species has been recognized as a species complex consisting of at least three
species. One of them, E. goeldlini Mazanek, Laska & Bicik, 1999, appears to be widespread in
Europe, and it has been found recently in all our neighbouring countries, but not in Belgium so
far. Given the presence in our neighbouring countries, it came as no surprise when we fïnally
found a female E. goeldlini in Belgian collection material. E. goeldlini can now also be added to
the Belgian fauna list.

Samenvatting. Eupeodes goeldlini (Diptera: Syiphidae) nieuw voor de Belgische fauna
Bij een recente revisie van het genus Eupeodes en van E. bucculatus (Rondani, 1857) in het
bijzonder is gebleken dat een aantal tot dan onopgemerkte soorten schuilgingen onder deze
soortnaam. Eén van deze soorten, Eupeodes goeldlini Mazanek, Laska & Bicik, 1999, heeft een
wijde Europese verspreiding en werd reeds in al onze buurlanden vastgesteld. Het was dan ook
geen verrassing dat bij de controle van Belgisch collectiemateriaal ook een vrouwtje Eupeodes
goedlini gevonden werd. Deze soort kan nu toegevoegd worden aan de Belgische faunalijst.

Résumé. Eupeodes goeldlini (Diptera: Syrphidae) espèce nouvelle pour la faune beige
Lors d'une révision récente du genre Eupeodes , et de E. bucculatus (Rondani, 1857) en
particulier, il apparait qu’il s'agit d'un groupe d'au moins trois espèces. Une d'entre elles, E.
goeldlini Mazanek, Laska & Bicik, 1999, est distribuée en Europe, et elle a été mentionnée de
tous nos pays avoisinants. II n'est donc pas surprenant que cette espèce a été trouvée aussi lors
d'une inspection des collections contenants des exemplaires beiges. E. goeldlini peut être ajouté a
la liste faunistique de Belgique.

Key words: Eupeodes goeldlini - Belgium - First record - faunistics.

Van de Meutter, F.: Achterheide 16, B-3980 Tessenderlo.

Frank.VandeMeutter@bio.kuleuven.be

Introduction

Eupeodes hoverflies are archetypical Syrphidae with yellowish transverse
banding or interrupted banding on the abdomen, which gives them a wasp-like
appearance. Due to a number of migrants and generalist species within this
genus, Eupeodes are a common member of Syrphidae assemblages throughout
Europe in all kinds of habitats. For Belgium, seven Eupeodes species have been
recorded so far. One of the species, E. bucculatus (Rondani, 1857) has a
tumultuous taxonomie history. This species was formerly known as E.
latilunulatus (Collin, 1931) or even before as Metasyrphus latilunulatus , and
keys out as such in Verlinden (1991). A recent revision by Mazanek et al. (1998)
revealed that a senior synonym existed and the official species name was
changed to E. bucculatus (Rondani, 1857). Soon after this, it appeared that
European E. bucculatus consisted of at least three different species: E.
bucculatus (Rondani, 1857), E. duseki Mazanek, Laska & Bicik, 1999, and E.
goeldlini Mazanek, Laska & Bicik, 1999. E. duseki so far is only known from
northem Scandinavia (Sweden, Norway; Speight 2007). The other two species,
however, are widely distributed over Europe, and have their range extending as
far as East-Russia and China. Eupeodes goeldlini appears to be not particularly

Phegea 37 (2) (01.VI.2009): 51

rare, and it has recently been reported from Britain, Czech Republic, Germany,
Finland, France, Ireland, the Netherlands, Poland, and Switzerland (Speight
2003, 2006). In the Netherlands, nearly half of the specimens of E. bucculatus
s.1. that have been checked tumed out to be E. goeldlini (Reemer & van Steenis
2006). As a consequence, this species was expected to be present in Belgium as
well.

Material and methods

While checking my personal Syrphidae collection I discovered a female
labelled as E. bucculatus that keyed out as E. goeldlini. This individual was
trapped on 28. V. 1998 at Dorstveld, Bonheiden, Prov. Antwerp, leg. & coll. Van
de Meutter Frank. This place is a 0.25 ha oldfield open area within deciduous
forest. Within 300 m range from this locale are extensive wet and dry meadows,
old parkland and Pinus coniferous stands. This high variety of habitats is due to
its location at the northem flank of the Dyle valley which exhibits a moisture
gradiënt from open water and marshland to forested dry sand dunes (former
heathlands) within a few hundred meters. This makes it difficult to derive any
habitat characteristics of E. goedllini from this record. Interestingly, a female E.
bucculatus trapped on 09.VI.1999 at exactly the same place is present in my
collection, indicating both species may co-occur.

The identification of Eupeodes can be troublesome. Eupeodes in general and
E. bucculatus in particular exhibit a high morphological variation (which has
given rise to its Dutch name "variabele kommazwever"), especially in the extent
and intensity of the banding pattems on the abdomen. I used the key in Speight
(2007) to re-identify all Eupeodes present in my collection. E. goeldlini keyed
out perfectly by having the lateral margin of tergite 5 entirely yellow (Fig. 1),
large dust spots on the frons (Fig. 2), the alula completely covered by
microtrichia (Fig. 2), stemite 4 with a rounded black mark and stemite 5 entirely
pale (Fig. 1), hind femora entirely yellow (Fig. 1), the posterolateral hair fringe
on the front femur was yellowish pale and the second basal cell of the wing had
less than 50% of its surface bare of microtrichia. All identification features could
be checked against female E. bucculatus , E. nitens and E. nielseni individuals in
my collection (all from Flanders) of which the identity could be confmned with
the key of Speigth (2007).

Phegea 37 (2) (01.VI.2009): 52

Fig. 1: Dorsal (left) and ventral (right) view on female Eupeodes goeldlini from Bonheiden
28.V.1998.

Fig. 2: Detail of the frons with dust markings (left) and alula with microtrichia (right) of the female
Eupeodes goeldlini from Bonheiden 28.V.1998.

Conclusion

This is the first report of the presence of E. goeldlini in Belgium, and this
species can be added to the Belgian fauna list. It is expected that E. goeldlini
occurs more widely throughout Belgium, as is the case in the Netherlands. So
far, there is no clear defmition neither of the habitat preferences of E. goeldlini,
nor whether these may differ from those of E. bucculatus. The fmdings of
Speight (2007) and Reemer & van Steenis (2006) contradict on whether one of
the species prefers more or less moist conditions. Our observations indicate that
both species may co-occur, and it is possible that habitat differences, if any, are
very subtle. At present, it appears that E. goeldlini may be expected in any of the
habitats where E. bucculatus s.1. has been found before.

E. bucculatus s.1. has a wide but scattered distribution throughout Belgium.
A relatively large number of observations were done since 1980 (64, Belgian
database). They appear to occur typically at low densities, for example, only
three records in the database had more than two individuals. Finding E.

Phegea 37 (2) (01.VI.2009): 53

bucculatus and allies requires searching for Eupeodes with slender abdominal
banding, although they may have separated spots as well. It is expected that
more E. goeldlini will turn up the next few years in Belgium, if they are better
looked for.

References

Mazanek, L., Bicik, V. & Laska, P. 1998. Redescription and reinstatement of Eupeodes bucculatus
(Rondani, 1857) and its synonymy (Dipt., Syrphidae). — Acta Universitatis Palackianae
Olomucensis Facultas Rerum Naturalium, Biologica 36: 27-38.

Mazanek, L., Laska, P. & Bicik, V. 1999. Two new Palaearctic species of Eupeodes similar to E.
bucculatus (Diptera, Syrphidae). — Volucella 4: 1-9.

Reemer, M. & van Steenis, J. 2006. Het zweefvliegduo Eupeodes bucculatus en E. goeldlini in
Nederland (Diptera: Syrphidae). — Nederlandse Faunistische Mededelingen 25: 87-92.

Speight, M. C. D. 2007. Species accounts of European Syrphidae (Diptera), Espoo, 2007. - In:
Speight, M. C. D., Castella, E., Sarthou, J.-P. and Monteil, C. (eds.) Syrph the Net, the database
of European Syrphidae 55, 286 pp. — Syrph the Net publications, Dublin.

Speight, M. C. D., Sarthou, J.-P. & Levy, D. A. 2007. Eupeodes goeldlini (Dip.: Syrphidae) new to
Britain, France and Ireland, with a key to separate it from related Atlantic zone species. —
Entomologist's record and Journal of Variation 1 19(5): 2 1 3-2 19.

van Veen, M. P. 2004. Hoverflies of Northwest Europe. — KNNV Uitgeverij, Utrecht.

Verlinden, L. 1991. Zweefvliegen (Syrphidae). — Koninklijk Belgisch Instituut voor
Natuurwetenschappen, Brussel.

Phegea 37 (2) (01.VI.2009): 54

Descriptions of new Pyraloidea from the Palaearctic
Region (Lepidoptera, Pyraloidea, Pyralidae:
Pyralinae, Phycitinae, Crambinae)

Jan Asselbergs

Abstract. Four new Pyraloidea species are described: Phycita characterica sp. n.
(Phycitinae), Bazaria venosella sp. n. (Phycitinae), Euzophera hulli sp. n. (Phycitinae), and Talis
caboensis sp. n. (Crambinae). Adults and genitalia are illustrated, distinctive characters for
sibling species are given. Scotomera caesarealis (Ragonot, 1891) (Pyralinae) is mentioned for
the fïrst time for the Continental European fauna.

Samenvatting. Beschrijving van nieuwe Pyraloidea uit het Palaearktische gebied
(Lepidoptera, Pyraloidea, Pyralidae: Pyralinae, Phycitinae, Crambinae)

Vier nieuwe soorten Pyraloidea worden beschreven: Phycita characterica sp. n. (Phycitinae),
Bazaria venosella sp. n. (Phycitinae), Euzophera hulli sp. n. (Phycitinae) en Talis caboensis sp.
n. (Crambinae). Adulten en genitalia worden afgebeeld, onderscheidingskenmerken voor
gelijkende soorten worden besproken. Scotomera caesarealis (Ragonot, 1891) (Pyralinae) wordt
voor het eerst uit continentaal Europa vermeld.

Résumé. Description de nouveaux Pyraloidea de la région paléarctique (Lepidoptera,
Pyraloidea, Pyralidae: Pyralinae, Phycitinae, Crambinae)

Quatre nouvelles espèces de Pyraloidea sont décrites: Phycita characterica sp. n. (Phycitinae),
Bazaria venosella sp. n. (Phycitinae), Euzophera hulli sp. n. (Phycitinae) et Talis caboensis sp.
n. (Crambinae). Les adultes et les genitalia sont figurés et des caractères diagnostiques sont
donnés. Scotomera caesarealis (Ragonot, 1891) (Pyralinae) est mentionné ici pour la première
fois de la région européenne continentale.

Zusammenfassung. Beschreibung von neue Pyraloidea aus dass Palaarktisches Gebiet
(Lepidoptera, Pyraloidea, Pyralidae: Pyralinae, Phycitinae, Crambinae)

Vier neue Pyraloidea Arten werden beschrieben und zusammen mit den Genitaliën abgebildet:
Phycita characterica sp. n. (Phycitinae), Bazaria venosella sp. n. (Phycitinae), Euzophera hulli
sp. n. (Phycitinae) und Talis caboensis sp. n. (Crambinae). Unterscheidungsmerkmale fiir
ahnliche Arten werden diskutiert. Neu fiir kontinentales Europa ist Scotomera caesarealis
(Ragonot, 1891) (Pyralinae).

Key words: Pyraloidea - Pyralinae - Phycitinae - Crambinae - new species.

Asselbergs, L: Neerland 20, NL-4614 GD Bergen op Zoom. The Netherlands.

jef.asselbergs@hetnet.nl

Introduction

The Pyraloidea form a very large superfamily with an estimated number of
16000 species worldwide. It is estimated that the same number of species is still
awaiting description. The current knowledge about the occurrence of Pyraloidea
species in Europe is summarized by Nuss, M., Segerer, A. & Speidel, W.
Although the pyraloid fauna of the Western Palaearctic region is relatively well
known, descriptions of new species from the lesser well known parts such as i.e.
Turkey, Southern Europe, the Canary Islands and Madeira are not infrequent. In
this paper a description is given of 4 new Phycitinae and 1 new Crambinae
species. Moreover, Scotomera caesarealis (Ragonot, 1891) is mentioned for the
first time from Continental Europe.

Phegea 37 (2) (01.VI.2009): 55

Figs. 1-10: 1 - Scotomera caesarealis (Ragonot, 1891) <$, Greece, Fokis, Itea, 20.vii.1987, leg. H. G.
van der Wolf; 3 - Phycita characterica sp. n., Holotype <$, Turkey, Antalya, 40 km N Alanya, 1350
m, ó.viii. 1 998, leg. W. De Prins; 5- Phycita mianella Amsel, 1949 S, Turkey, Mersin,10 km S
Arslanköy, Taurus, 1300 m, ll.vii.1987, leg. M. Fibiger; 7 - Bazaria venosella sp. n., Holotype S,
Canary Islands, La Palma, Tacande, ll.iii.2007, leg. H. W. van der Wolf; 8 - B. venosella sp. n.,
Paratype S, Malta, Naxxar, 7.iv.l998, leg. A. Seguna; 9- B. venosella sp. n., Paratype c^,Tenerife,
Tamaimo,13-23.iii.l999, leg. M. Delnoye; 10 - B. venosella sp. n., Paratype Canary Islands,
Tenerife, Los Christianos, 10-20.iii.1980, leg. J. B. Wolschrijn.

Material and methods

An Olympus stereo microscope type VT-II has been used for the descriptions
of the adults. Genital slides were made with a Beek microscope type CBS and
the drawings of the genitalia were made with an Olympus microscope CH2 with
a drawing tube. Terminology for the morphological structures follow Roesler,
(1973, 1993). If not stated otherwise, measurements of the forewing include the
fringe.

Phegea 37 (2) (01.VI.2009): 56

Abbreviations:

coll. collection

gen. prep. genital slide preparation

ZMFK Zoologisches Forschungsinstitut und Museum Alexander Koenig , Bonn , Germany.

IEEM Instituto Espanol di Entomologia , Madrid, Spain.

LPM Liverpool Museum , Merseyside , U.K.

ZMA Zoologisch Museum , Amsterdam , The Netherlands.

BMNE1 British Museum of Natural History, London, U.K.

Scotomera caesarealis (Ragonot, 1891) (Pyralinae) (Figs. 1, 2) new to the
Continental European fauna.

Material: 1 S, Greece, Fokis, Itea, 20.vii.1987, leg. H. G. van der Wolf, gen. prep. 2736,
Asselbergs, coll. Asselbergs.

Remark. The species was identified by M. Shaffer, BMNH.

Life history. Unknown.

Distribution. Known from N. Syria [present-day Turkey], Marasch
(Osthelder 1935); Palestine, Jordan Valley (Zemy 1914); Palaearctic Egypt
(Rebel 1939); Europe: Cyprus, Limassol (Rebel 1939); mainland of Greece:
Fokis, Itea, (this paper).

Phycita characterica sp. n. (Phycitinae) (Figs. 3, 4)

Material. Elolotype S, Turkey, Antalya, 40 km N Alanya, 1350 m, ó.viii. 1 998, leg. W. De Prins,
gen. prep. 4313 Asselbergs, ZMA. Paratype S, same data, gen. prep. 4371 Asselbergs, coll.
Asselbergs.

Description. Forewing length 12.5 mm; alar expanse 27 mm. Head: vertex
flatly rounded, covered with adjacent scales. Ocelli and chaemosemata present.
Labial palpi 1.5x eye, slender and uptumed, 3rd segment straight, 0.5x 2nd
segment. Maxillary palpi filiform, 0.25 x 2nd segment of labial palpi. Patagium
greyish brown as the thorax, necktuft with raised scales. Antenna with a strong
sinus and a distinct dark and shiny scalebush, flagellum ciliated. Forewing
greyish brown almost without a pattem, except for the postmedian line which
originates from the costa at 4/5 and is next running with an outward curve to the
inner margin near the tomus. Fringe greyish brown with a white base and a
white dividing line. Flindwing pale greyish brown, little hyaline. Base of fringe
white followed by a brown dividing line , next greyish.

Male Genitalia: Uncus subtriangular , 1.2x longer than broad, base flat,
apex flattened. Gnathos triangular, sharply tapering towards a curved apex, the
total length being slightly less than 0.25 x uncus. Scaphium rectangular, 5/8 x as
long as the uncus; lateral gnathos branches oblique, proximally with extensions.
Transtilla elements slightly less than 1.5x gnathos, elongate and narrowly
triangular. Anellus U-shaped with sclerotized base and slender digitate processes
which are distally provided with a few setae. Valva slender, slightly widening
towards a rounded cucullus; costa proximally with a globular lump, followed by
a semi circular projection and a costal enforcement, ending abruptly and sharply
pointed proximally from the cucullus. Vinculum V-shaped, 1.25x longer than

Phegea 37 (2) (01.VI.2009): 57

valve, base flat. Length of phallus slightly less than 1.3* valve and 4/5 x longer
than broad. Vesica with a conspicuous big and curved comutus with an
extended base and with a curved part well extending outside the phallus wall; in
the middle of the curved part originates a smaller and distally slightly dentate
comutus also extending beyond the phallus wall. Base of culcita centrally
concave, at both sides first a short scale bundle of which 1 scale is 2x longer
than the others, followed distally by a scale bundle which is 2x longer than the
valve.

Fig. 2. Scotomera caesarealis (Ragonot, 1891), male genitalia; a - genitalia; b - aedeagus; c-
everted vesica with comuti.

Phegea 37 (2) (01.VI.2009): 58

Fig. 4. Phycita characterica sp. n.; a - genitalia; b- aedeagus; c - culcita.

Differential diagnosis. Extemally, Phycita characterica is similar to P.
mianella Amsel, 1949 (Figs. 5, 6), but the male genitalia are quite different: P.
characterica has an elongate subtriangular uncus (broadly ovate in P. mianella );
In P. characterica the transtilla elements are slender, basally not expanded
(stouter, longer and basally expanded in P. mianella ); presence of costal
projections in P. characterica (absent in P. mianella ); phallus with a big curved

Phegea 37 (2) (01.VI.2009): 59

comutus in P. characterica (1 straight comutus and a bundle of long slender
comuti in P. mianella).

Derivatio nominis. Named after the conspicuous comutus in the vesica.
Life hi story. Unknown.

Distribution: Only known from the type locality, Turkey, Antalya, 40 km
N Alanya.

Fig. 6. Phycita mianella Amsel, 1949; a - genitalia; b. aedeagus; c - culcita.

Phegea 37 (2) (01.VI.2009): 60

Bazaria venosella sp. n. (Phycitinae) (Figs. 7, 8, 9, 10)

Material. Holotype S, Canary Islands, La Palma, Tacande, ll.iii.2007, leg. H. W. van der Wolf,
gen. prep. 5835 Asselbergs, ZMA. Paratypes, 1$, Canary Islands, Tenerife, Los Christianos, 10-
20.iii. 1 980, leg. J. B. Wolschrijn, gen. prep. 3917 Asselbergs, coll. Asselbergs; 1(7, Tenerife,
Tamaimo, 13-23. iii. 1999, leg. M. Delnoye, gen. prep. 4393 Asselbergs, coll. Cox; 1(7, Malta,
Naxxar, 7.iv.l998, leg. A. Seguna, gen. prep. 5793 Asselbergs, coll. Seguna.

Description. Alar expanse 19-23 mm, length of forewing 8.5-10.5 mm
(males), resp. 21.5 and 10 mm (female). Male: vertex with cream coloured scales
surpassing frontal edge of eye with 1/3 eye diameter. Ocelli and chaemosemata
present. Labial palps 1.75x eye, cream coloured with a few brown scales at the
outside; 3rd segment 0.25 * 2nd segment and slightly drooping. Maxillary palps
hair-pencil shaped and hidden in a groove of the labial palps. Proboscis normally
developed. Scape of antennae 2/5 longer than broad. Sinus flat, segments 3-8
with tiny sclerified thoms, flagellomers pubescent. Patagia and thorax pale
ochreous. Forewing pale ochreous, more or less sprinkled with brown scales
along the veins. Fringe as the ground colour, i.e. whitish ochreous till white.
Hindwing whitish, fringe white. Female: Antennas without sinus and
sclerifications, pubescent. Maxillairy palps small, flat and adjacent to frons.
Hindwing pale ochreous, a little hyaline and with a terminal brown line. Fringe
brown at base, next snow white.

Fig. 1 1. Bazaria venosella sp. n.; a.- genitalia, b. aedeagus; c.- culcita.

Phegea 37 (2) (01.VI.2009): 61

Male Genitalia: uncus triangular, base and apex flattened, greatest width
about 7/9 of length, bilaterally with a few hair-like setae. Gnathos 0.5 x uncus,
ovate. Scaphium not depassing apex of uncus. Tegumen slender. Transtilla
components slender, about 4/5 x gnathos; they are weakly sclerotized and
sometimes invisible. Anellus V-shaped, processes withhout hair-like setae.
Vinculum almost parallel-sided till slightly V-shaped, base flatly rounded. Valva
with straight costa, cucullus rounded. Sacculus distally with a sclerified
triangular process. Valve below costa with a long sclerotized tapering bar
extending till just beyond cucullus. Phallus stout, about 2.3 x longer than broad.
Vesica with a longitudinal band of cornuti of different size, 2/3 of phallus
length. Culcita bilateral with 5 bundles of differently shaped scales, the longest
having slightly less than 1.5x valve length.

Fe male genitalia: Papillae anales triangular, about 1.5X longer than
broad, 8th segment 2/3 x longer than broad. Posterior apophyses almost reaching
proximal edge of 8th segment and about as long as anterior apophyses. Length of
ductus bursae l/7x bursa, proximally scobinate. Bursa about 3x longer than
wide and distally with 2 rows of oppositely directed cornuti; the middle part of
the bursa has a curved band of cornuti, distally of this band originates the ductus
seminalis. The proximal part of the bursa is slightly wrinkled.

Fig. 12. Bazaria venosella sp. n., female genitalia.

Phegea 37 (2) (01.VI.2009): 62

Differential diagnosis. Extemally, specimens of Bazaria venosella sp.
n. with marked veins are not unsimilar to Ancylosis nervosella (Zemy, 1914), or
to Ancylosis sulcatella (Christoph, 1877). The male genitalia resemble those of
Bazaria leucochrella (Herrich-Schaffer, 1852), viz. Roesler (1993). In B.
leucochrella the costal enforcement is only partly free from the costa (free from
the costa in B. venosella); the vesica has 1 long comutus in B. leucochrella (a
longitudinal band of numerous small comuti in B. venosella ).

Derivatio nominis. Named after the more or less brownish suffusion
along the veins.

Life hi story. Unknown. Specimens were caught at light in March and
April.

Distribution. Canary Islands: La Palma: Tacande; Tenerife: Los
Christianos, Tamaimo; Malta: Naxxar.

Euzophera hulli sp. n. (Phycitinae) (Figs. 13, 14, 15, 16)

Material. Holotype S, Turkey, Kayseri, Sultansazligi, 1183 m, 4.VÜ.1995, leg. M. Huil, gen. prep.
5171 Asselbergs, LPM; Paratypes 2<$, gen. prep. 5177 and 5180Asselbergs, 1$, gen. prep. 5178
Asselbergs, all same data, coll. Huil and coll. Asselbergs.

Description of the male. Forewing 13-14.5 mm; alar expanse 27-32
mm. Head: vertex flatly rounded. Labial palpi porrect, 2.25 x the eye, cream
coloured at the inside and reddish brown at the outside. Maxillairy palpi 0.8x 3rd
segment of labial palpi. Ocelli invisible, chaetosemata present. Proboscis
strongly reduced. Scape 1.5 longer than broad, sinus absent. Male flagellum uni-
pectinate. Patagium, tegulae and forewing yellowish ochreous with a few reddish
scales. Antemedian line from the costa at 1/5 to the inner margin at 1/4 and only
visible by a few reddish scales at the outer border of the line. Postmedian line
from the costa at 5/6 to the inner margin at 4/5, the line is faintly edged with
reddish scales at the inside. There are 2 brown discal spots, the one most near to
the costa slightly displaced in the direction of the wingbase. Fringe yellowish
white. Hindwing whitish, dusted greyish brown, except for the costa, the
wingbase and the inner margin which are white.

Fe male: as the male but with pubescent antennae. Forewing with some
reddish suffusion.

Male genitalia: Uncus an almost equilateral triangle, apex rounded.
Gnathos straight and parallel sided, apically slightly narrower and with a small
cleft, staying below distal margin of uncus. Lateral gnathos branches without
particularities. Transtilla medially fused, the components expanded proximally.
Tegumen slender. Juxta U-shaped, digitate processes apically with a few hair-
like setae. Length of valve slightly less than 3x width, distally from sacculus
abruptly widening; costal enforcement along the whole of the costa.

Female genitalia: Ovipositor triangular, slightly longer than broad.
Length of 8th segment about %x width. Length of posterior apophyses about 4/5
of anterior apophyses. Ductus bursae 3/5 of corpus bursae with in the buiging

Phegea 37 (2) (01.VI.2009): 63

middle part a group of granulate sclerifications. Bursa ovate with a group of tiny
granules just below the origin of the ductus seminalis in the distal part of the
bursa.

Fig. 14. Euzophera hulli sp. n.; a - genitalia; b- aedeagus; c - culcita.

Vinculum proximally rounded and slightly more than 2x broader than long.
Phallus 1.7x valve and 5x longer than broad. Vesica terminally with 2 bundles
of tiny comuti. Culcita simple with a flatly rounded basal list and bilateral with a
short scale-bundle of about l/3x valva.

Remark. Although the male and the female genitalia are of the Euzophera
type, the long and porrect labial palps and the unipectinate antennas in the male
are uncommon characters for Euzophera , in which hulli sp. n. is placed
provisionally.

Differential diagnosis. At present, none of the known Euzophera
species could be confounded with E. hulli; superficially there is some similarity
with E. kabulella Roesler, 1973 which is smaller with a wingspan of 17 mm,
with labial palps of 1/3 x the eye in the male and with a pubescent flagellum.

Derivatio nominis. Named in honour of Michael Huil , who collected the
new species.

Life history. Unknown.

Distribution. Turkey: Kayseri, Sultansazhgi, 1183 m; probably it has a
wider distribution.

Phegea 37 (2) (0 1 . VI.2009): 64

Figs. 13-19: 13 - Euzophera hulli sp. n., Holotype S, Turkey, Kayseri, Sultansazhgi, 1183 m,
4.VÜ.1995, leg. M. Huil; 15- E. hulli sp. n., Paratype $, Turkey, Kayseri, Sultansazhgi,
4.vi. 1995, leg. M. Huil; 17 - Talïs caboensis sp. n., Holotype S, Spain, Almeria, Cabo de Gata, 11-
13.X.2007, leg. A. Cox; 19 - T. caboensis sp. n., Paratype $, same data.

Talis caboensis sp. n. (Crambinae) (Figs. 17, 18, 19, 20)

Material. Holotype S, Spain, Almeria, Cabo de Gata, 1 1-13.X.2007, leg. A. Cox, gen. prep. 5338
Asselbergs, ZMA. Paratypes, 94c?, 1$, gen. prep. 5852 Asselbergs; 1$, same data, coll. Cox, 3$,
same data, coll. Asselbergs; lc?, Spain, Almeria, Cabo de Gata, 21.X.2003, leg. Viehmann, gen. prep.
5536 Asselbergs, coll. Schmitz.

Description. Male: alar expanse 26.5 mm, forewing 12.5 mm. Ocelli and
chaemosemata present. Frons strongly porrect, triangular, apex pointed; head
and frons covered with brownish scales, at both sides with cream coloured
scales. Patagia, tegulae and thorax brownish as the forewings. Proboscis
reduced. Labial palpi porrect, 3x the eye, cream coloured at the inside, brown at
the outside. Maxillary palpi long and pencil-like, proximal half brown, distal half
cream coloured. Proboscis reduced. Frenulum as a long thom in male and
female. Male antennas unipectinate with narrow rhomboid base. Forewings dull
brown, sparsely sprinkled with a few half erected black scales. Antemedian line
from the costa at 1/2 with 2 outwardly directed angles to dorsum at 2/5.
Postmedian line from the costa at 6/7, partly whitish, outwardly convex and with
an inwardly directed angle above dorsum to 5/6 on dorsum. Discocellular spot
prominent, whitish. Fringe brownish, base white. Hindwings slightly shiny grey-
brown. Fringe whitish divided by a greyish line. Female as the male but with
filiform antennas.

Male genitalia: Uncus parallel-sided, apex pointed. Gnathos distally
drop-shaped. Valva gradually narrowing towards a rounded cucullus. Ventral

Phegea 37 (2) (01.VI.2009): 65

margin of valva with a blunt vault. Aedeagus parallel- sided, about as long as
valva. Vesica with 1 long slender comutus slightly longer than half of aedeagus.

Female genitalia: Ovipositor triangular. Apophyses posteriores stout,
depassing proximal edge of 8tfi segment with plm 1/7. Eight segment ± 5/3 x
broader than high. Ductus bursae relatively short, gradually transitioning into
corpus bursae. Corpus bursae stretched ovoid, distal part wrinkled.

Differential diagnosis. Nearest to Talis arenella Ragonot, 1887 and
Tal is afra (Bethune-Baker, 1894), which is known from Egypt, Alexandria, the
differences between these three species are shown in the table below:

Phegea 37 (2) (0 1 . VI.2009): 66

Fig. 18. Talis caboensis sp.n., male genitalia.

Fig. 20. Talis caboensis sp. n.,
female genitalia.

T. arenella

T. afra

T. caboensis sp. n.

exp. 28-29 mm (males)

exp. 21 mm (males)

exp. 26.5 mm males, 21 mm
(females)

valva ventrally with well-
defïned, rather slender
projection

valva ventrally without
projection

valva outwardly curved

comutus in vesica broader

comutus in vesica slender,
slightly bent in middle

comutus in aedeagus very
slender, at base wide, straight

Phegea 37 (2) (01.VI.2009): 67

The females of arenella and af ra are unknown.

Derivatio nominis. Named after the type locality .

Life hi story. Unknown.

Distribution. So far only known from Cabo de Gata in South Spain.

Acknowledgements

For the gift or loan of material the author is much indebted to the following
persons: Mr A. Cox (Mook, The Netherlands), Mr M. Huil (Gwynnedd, Wales,
UK), Mr W. De Prins (Leefdaal, Belgium), Mr W. Schmitz (Bergisch-
Gladbach, Germany), Mr A. Seguna (Naxxar, Malta), Mr H. W. van der Wolf
(Nuenen, The Netherlands), Mr J. B. Wolschrijn (Twello, The Netherlands), and
Mr M. Shaffer (BMNH, London, UK) for the identification of Scotomera
caesarealis.

References

Amsel, H. G. 1949. Die Microlepidopteren der Brandt'schen Iran-Ausbeute. I.Teil. — Bulletin de la
Société Fouadl. Entomologie 33: 227-269, 4 Taf.

Bleszynski, S. 1965. Pyralidae, Crambinae. - In: Amsel, H. G., Gregor, F. & Reisser, E. (eds.):
Microlepidoptera Palaearctica I. Crambinae. — Verlag Georg Fromme & Co., Wien, Textband
pp. 1-553, Tafelband Taf. 1-133.

Nuss, M., Segerer, A. & Speidel, W. 2004. Crambidae, Pyralidae, Thyrididae. - In: Karsholt, O. &
Nieukerken, E. J. van. Fauna Europaea: Lepidoptera, Moths. — Fauna Europaea version 1.1.,
http:// www.faunaeur.org.

Osthelder, L. 1935. Lepidopteren-Fauna von Marasch in turkisch Nordsyrien. — Mitteilungen der
münchener entomologische Gesellschaft 25: 56-58, 67-90.

Ragonot, E. L. 1891. Essai sur la classifïcation des Pyralites. Note supplementaire et rectifïcative. —
Annales de la Société entomologique de France 11: 599-662.

Rebel, H. 1939. Zur Lepidopterenfauna Zypems. — Mitteilungen der münchener entomologische
Gesellschaft 29: 487-564, Taf. 15.

Roesler, R. U. 1973. Pyralidae, Trifine Acrobasiina. - In: Amsel, H. G., Gregor, F. & Reisser, E.
(eds.) Microlepidoptera Palaearctica Band 4. Trifine Acrobasiina. — Verlag Georg Fromme &
Co., Wien. Textband pp. 1-752, Tafelband Taf. 1-170.

Roesler, R. U. 1993. Pyralidae, Quadrifme Acrobasiina. - In: Amsel, H. G., Gregor, F. & Reisser, E.
(eds.) Microlepidoptera Palaearctica Band 8. Quadrifme Acrobasiina. G. Braun Druckerei und
Verlage, Karsruhe. Textband pp 1-305, Tafelband Taf. 1-82.

Zemy, H. 1914. Über Palaearktische Pyraliden des K.K. Naturhistorischen Hofmuseums in Wien. —
Annalen des naturhistorischen Museum zu Wien 28: 295-348, Taf. 15-16.

Phegea 37 (2) (01.VI.2009): 68

New records of the rare Escher’s Blue, Polyommatus
escheri (Lepidoptera: Lycaenidae), from the Republic
of Macedonia

Micevski Branko, Micevski Nikola & Verovnik Rudi

Abstract. Polyommatus escheri (Hübner, 1823) was known only from a single site in
Republic of Macedonia. During field surveys in June 2008 the species was discovered at two
new sites. In both cases the habitat is characterised by stretches of barren sandy ground where the
larval food plant Astragalus monspesullanus (Bemh.) grows abundantly. The habitat is at least
partially anthropogenic and mainly consists of 100 to 300 m long road verges. Given the limited
habitat available at all three known sites and potential threats due to isolation, natural succession
and anthropogenic pressure, the species should be considered highly threatened in Republic of
Macedonia.

Rezime. Eseroviot sinec bese poznat samo od eden lokalitet za Republika Makedonija. Za
vreme na terenskite istrazuvanja vo juni 2008 vidot bese otkrien na uste dva novi lokaliteti. Vo
site tri slucai stanisteto se karakterizira so ogolena pesokliva podloga so znacitelna abundancija
na Astragalus monspesullanus (Bemh.), rastenie domakin za ovoj vid. Stanistata na site lokaliteti
se odlikuvat so delumna antropogenost i glavno se sostojat od pojasi od po 100-300 m kraj
samiot pat. Zaradi ogranicenosta na stanistata i potencijalnata opasnot kako rezultat na prirodna
sukcesija i antropogeniot impakt, vidot treba da se smeta za visoko zagrozen vo Makedonija.

Samenvatting. Nieuwe gegevens over Polyommatus escheri in de Republiek Macedonië.
Polyommatus escheri (Hübner, 1 823) was slechts van één lokaliteit in de Republiek Macedonië
bekend. Tijdens veldonderzoek in juli 2008 werd de soort in twee nieuwe vindplaatsen
aangetroffen. In beide gevallen gaat het om een stuk zandgrond waar de voedselplant van de
mps, Astragalus monspessulanus (Bemh.) overvloedig groeit. De vindplaatsen zijn op zijn minst
gedeeltelijk van anthropogene oorsprong en bestaan hoofdzakelijk uit wegboorden van 110 a 300
m lang. Gezien de kleine oppervlakte van de drie vindplaatsen en de mogelijke dreiging van
isolering en anthropogene dmk, moet de soort in de Republiek Macedonië beschouwd worden
als zeer bedreigd.

Résumé. Nouvelles observations de Polyommatus escheri en République de Macédoine.
Polyommatus escheri (Hübner, 1823) est connu seulement d'une localité en République de
Macédoine. Pendant des recherches en juillet 2008 cette espèce fut trouvée dans deux nouvelles
localités. Toutes les deux des endroits sablonneux oü la plante nourricière de la chenille,
Astragalus monspessulanus (Bemh.), croït abondamment. Ces localités sont, au moins en partie,
d'origine anthropique et consistent surtout en bords de route de 110 a 300 m de long. Vu la
superficie négligeable des trois localités connues et la pression humaine, P. escheri doit être
considéré comme extrêmement vulnérable en République de Macédoine.

Key words. Polyommatus escheri - Distribution - Faunistics - Republic of Macedonia.
Micevski, B. & Micevski, N.: Faculty of Natural Sciences, Institute of Biology, BSPSM, Gazi

Baba b.b., Skopje, Republic of Macedonia.

Verovnik, R.: University of Ljubljana, Biotechnical Faculty, Department of Biology, Veena pot

111, 1000 Ljubljana, Slovenia. mdi.verovnik@bf.uni-lj.si.

Introduction

The Escher's Blue, Polyommatus escheri (Hübner, 1823), is a typical
Mediterranean species distributed from Spain to NW Greece with isolated
populations in the Southern Alps and the Atlas Mountains in Morocco (Pro
Natura - SBN 1994, Tolman & Lewington 1997, Kudma 2002). In the Balkan

Phegea 37 (2) (01.VI.2009): 69

Peninsula it is present along the Adriatic coast, excluding Montenegro (Sijaric
1984), most parts of Greece (Pamperis 1997), SW part of Bulgaria (Abadijev
2001) and few isolated populations in Southern and eastem Serbia (Jaksic 1988,
Swaay et al. 2007). Given the relatively wide distribution of the species in the
region, especially northem Greece, it is hard to explain why it was found only at
a single site in Macedonia despite concerted effort during surveys for the
distribution atlas (Schaider & Jaksic 1989). The species is first mentioned for
Macedonia by Thumer (1964) who does not provide exact locality for the
species, other than near the village Drenovo. This agrees well with the site in
Raec Gorge about 3 km NW from Drenovo, where the species was collected by
Jaksic (Schaider & Jaksic 1989).

The species is a habitat specialist limited by the presence of larval food plant
growing mainly in pioneer associations (Pro Natura - SBN 1994, Verovnik
2004). Escher's Blue is monophagous utilizing a few species of Astragalus,
principally A. monspesullanus. This plant grows in open, sandy places, open
canopy forests, and forest edges. As such habitats are not rare in Macedonia the
species was expected to be found at new sites.

Field survey

During 31.5.-6.6.2008 a joint survey was undertaken by the authors visiting
many less studied areas in Macedonia for basic faunistic research. On 4.6.2008
the Raec gorge was visited and the larval food plant was present along a 50 m
long stretch of an abandoned road. Due to unfavorable weather conditions no
adults were observed. The habitat is very limited at the site with only few A.
monspesullanus present.

The second site was discovered in the middle part of the Treska valley on
6.6.2008. The larval food plant was present in a small side valley just south of
the beginning of the artificial lake Kozjak, around 3 km south of the village
Tazhevo and with NE exposition. It was growing along 300 m of a road verge
with some plants present in low density also on the steep slopes above it.
Butterflies of both sexes were completely fresh and present along the whole
stretch of the suitable habitat. In a single count approximately 15 males and 4
females were seen. The site was rich in butterflies with Carcharodus lavatherae
(Esper, 1783), Euchloe penia (Freyer, 1851), Iolana iolas (Ochsenheimer,
1816), and Scolitantides orion (Pallas, 1771) being the most prominent
examples. The exact position of the site is 41°41,838 N and 21°13,754 E at
altitude of 485 m.

Encouraged by the new fmding the first author surveyed the literature for
further records of A. monspesullanus in Macedonia. Surprisingly the species is
known only from 3 sites (Micevski 2001), one of which is in close proximity to
the newly recorded population. The only further promising site was vicinity of
Majdan (Kozuf Mts.) which was visited on 14.6.2008. The locality was similar

Phegea 37 (2) (01.VI.2009): 70

to the previous two, along the unsurfaced road, with SE exposition and near the
Bistra river.

Knowing the habitat, the site was easy to spot and both larval food plants and
adults were observed there. The site is situated along 300 m of road, few km
before the village Majdan. Here the adults were similarly abundant as in the
Treska valley. Further interesting records from this locality are Apatura ilia
(Denis & Schiffermüller, 1775), Euphydryas aurinia (Rottemburg, 1775),
Melitaea arduinna (Esper, 1783) and Erebia medusa (Fabricius, 1787). The
exact position of the site is 41°09,980 N and 21°56,810 E at altitude of 740 m.
Despite intensive search no further sites with larval food plant were found in the
vicinity.

Subspecific status

The specimens of Polyommatus escheri from Macedonia clearly belong to
the subspecies dalmatica (Speyer, 1882). The subspecies is characterized by the
broad marginal black border and bright blue coloration in the males.

Discussion

The new discoveries of Escher's Blue give hope that with further, even more
focused field work, the species could be discovered at new sites. This is
especially true for the entire Treska valley where numerous small side valleys
and steep slopes could be the potential main distribution area of the species in
Macedonia. Flowever, given the limited known range of both the larval food
plant A. monspesullanus (Micevski 2001) and P. escheri in Macedonia and their
ecological specialization the species should be considered highly threatened. If
we presume that all three sites where P. escheri is present are entirely isolated,
the long term survivor of these populations even without habitat changes is
unlikely. Both overgrowing due to succession and anthropogenic pressure
(destruction of road verges) could be potentially deleterious to these small and
isolated populations. Further research should focus on Treska valley where
despite large scale inundation the habitats for the species could still be present
and could provide a wider and thus less sensitive network of habitat patches.

Acknowledgments

Authors would like to thank Angel Keymeulen, Milan Buric, Chris van
Swaay, Karls Veling and Svetlana Miteva for their company during field work
and their valuable observations.

Literature

Abadjiev, S. P. 2001 . Art atlas of the distribution of the butterflies in Bulgaria. — Pensoft Publishers,

Sofia, 335 pp.

Jaksic, P. 1988. Provisional distribution maps of the butterflies of Yugoslavia. — Acta Entomologica

Jugoslavica - editiones separata I: 1-214.

Kudma, O. 2002. The Distribution Atlas of European Butterflies. — Oedippus 20: 1-344.

Micevski, K. 2001. The Flora of the Republic of Macedonia. — Macedonian Academy of Sciences 5:

1121-1430.

Phegea 37 (2) (01.VI.2009): 71

" ' •• *

T

&J. 4

as»

«4,

. : \ :

'M

Fig 1. Localities of Polyommatus escheri (Hübner, 1823) and its food plant A. monspesullanus
(Bemh.) in the Republic of Macedonia. The empty spot denotes the presence of the food plant only.
1. - Makedonski Brod, between Samokov and Kalugerec villages (Micevski 2001); 2. - Treska
valley, small side valley at the Southern edge of the artificial lake Kozjak, 3 km south of the village
Tazhevo; 3. - Kavadarci, Raec gorge 3 km NW from Drenovo (Thumer 1964, Schaider & Jaksic
1989); 4. - Kozuf Mts., along the road in Bistra river valley before the village Majdan.

Figs. 2-5. Polyommatus escheri (Hübner, 1823). 2 - Male upperside, Tazhevo near Kozjak Lake; 3-
Female underside, idem; 4 - Larval foodplant, Astragalus monspessulanus, idem; 5 - Locality near
Majdan village in Kozuf Mts. where adults of P. escheri were observed along the road side.

Phegea 37 (2) (01.VI.2009): 72

Pamperis, L. N. 1997. The Butterflies of Greece. — A. Bastas-D. Plessas Graphic Arts S.A., Athens,
XII + 559 pp.

Pro Natura - Schweizerischer Bund für Naturschutz (SBN), 1994. Schmetterlinge und ihre
Lebensraume, Arten-Gefahrdung-Schutz. Vol 1. — Schweizerischer Bund für Naturschutz,
Basel, 516 pp.

Schaider, P. & Jaksic, P. 1989. Die Tagfalter von Jugoslawisch Mazedonien (Rhopalocera und
Hesperidae). — Selbstverlag Paul Scheider, München, 199 pp.

Sijaric, R. 1984. The investigation of Rhopalocera (Lepidoptera in Montenegro). — The Montenegrin
Academy of Sciences and Arts Glasnik of the section of natural Sciences 4: 163-175.

Swaay C. van, Jaksic P. & Buric M. 2007. Notes on some summer butterflies (Lepidoptera:
Hesperioidea & Papilionoidea) of Eastem Serbia. — Acta entomologica serbica 12 : 1-10.

Thumer, J. 1964. Die Lepidopterenfauna jugoslavisch Mazedoniens. I. Rhopalocera, Grypocera und
Noctuidae. — Posebno Izdanie. Prirodonaucen Muzej Skopje 1: 1-158.

Tolman, T. & Lewington, R. 1997. Collins field guide. Butterflies of Britain and Europe. — Harper
Collins Publishers, London, 320 pp.

Verovnik, R. 2004. Distribution and conservation status of Polyommatus escheri (Huebner, 1823) in
Slovenia (Lepidoptera: Lycaenidae). — Linneana Belgica 19: 253-257.

Phegea 37 (2) (01.VI.2009): 73

Nomenclature changes in Afrotropical Pterophoridae
(Lepidoptera)

Cees Gielis

Gielis, C.: Mr. Haafkensstraat 36, 4128 CJ Lexmond, the Netherlands. C.Gielis@net.hcc.nl.

Early 2008 (Gielis, 2008) I have described some new species of
Pterophoridae from the African continent. Recently I have visited the Transvaal
Museum in Pretoria, Republic of South Africa. This collection holds a great
number of specimens from the Southern half of Africa. It was here I discovered
that I had created two synonyms. Also the generic position of the valid species
needed to be changed. The results are tabulated as:

Pselnophorus jaechi (Arenberger, 1993) ( Pterophorus ), Comb. n.

Type locality: Kenya.

Pselnophorus meruensis Gielis, 2008, Syn. n.

Type locality: Tanzania.

Eucapperia bullifera (Meyrick, 1918) (. Platyptilia ), Comb. n.
Type locality: Republic of South Africa.

Eucapperia continentalis Gielis, 2008, Syn. n.

Type locality: Tanzania.

Reference

Gielis, C. 2008. Ten new species of Afrotropical Pterophoridae. — Zoologische Mededelingen,
Leiden 82: 43-57.

Phegea 37 (2) (01.VI.2009): 74

Nieuwe vondsten van prachtkevers aan de westrand
van Brussel (Coleoptera: Buprestidae)

Willy Troukens

Abstract. New records of Buprestidae (Coleoptera) at the westside of Brussels, Belgium.

In 2007, fïve interesting species of Buprestidae were found at the westside of Brussels: Anthaxia
nitidula (Linnaeus), Agrilus cyanescens Ratzeburg, A. sulcicollis Lacordaire, A. biguttatus
(Fabricius), and Buprestis novemmaculata Linnaeus. The fifth species was found in the
collections of the K.B.I.N., Brussels. It is apparently an adventive specimen.

Résumé. Nouvelles découvertes de buprestes a la périphérie ouest de Bruxelles, Belgique
(Coleoptera: Buprestidae).

En 2007 cinq espèces intéressantes de buprestes furent observées a la périphérie ouest de
Bruxelles: Anthaxia nitidula (Linnaeus), Agrilus cyanescens Ratzeburg, A. sulcicollis Lacordaire,
A. biguttatus (Fabricius) et Buprestis novemmaculata Linnaeus. La cinquième espèce a été
retrouvée dans la collection de E. Derenne qui se trouve actuellement a 1'Institut royal des
Sciences naturelles de Belgique. II s'agit manifestement d'un coléoptère adventice.

Key words: Belgium - faunistics - Buprestidae - Anthaxia nitidula - Agriclus cyanescens -
Agrilus sulcicollis - Agrilus biguttatus - Buprestis novemmaculata - Coleoptera.

Troukens, W.: Ninoofsesteenweg 782/8, B-1070 Anderlecht.

In dit tijdschrift verschenen eerder al enkele artikels over prachtkevers
(Buprestidae) aan de westrand van Brussel (Troukens 2005: 76-79, 2008: 101—
102). Maar de natuur geeft slechts met mondjesmaat haar geheimen prijs.
Onafgebroken veldwerk leidt daarom steeds weer tot nieuwe, verrassende
ontdekkingen.

Anthaxia nitidula (Linnaeus, 1758) (Bloemenprachtkever)

Remi Guinez nam op 24 mei 2007 tal van groenglanzende exemplaren van
het bloemenprachtkevertje, Anthaxia nitidula , waar op een bloeiende
egelantierstruik {Rosa rubiginosa ) die naast enkele volkstuintjes groeide te Vorst
(Brussel). In Brabant is dit prachtkevertje uiterst zeldzaam. Ikzelf vond ooit een
tiental exemplaren op een stervende pruimelaar {Prunus sp.) te Dilbeek
(Troukens 2005: 76-79),

Agrilus cyanescens Ratzeburg, 1837 (Blauw smalbuikje) (fig. 1)

Op 4 juni 2007 nam Remi Guinez eveneens te Vorst een staalblauw
prachtkevertje waar dat wij in de streek nog nooit hadden gezien. Het bleek te
gaan om het blauw smalbuikje, Agrilus cyanescens , een kleine soort van 4,5 a
7,5 mm. De kop en het halsschild zijn gerimpeld en de dekschilden zijn aan de
basis stevig ingedeukt. Volgens Keer (1930: 740) kan men de imago's in juni en
juli aantreffen op bloemen. De larven ontwikkelen zich in verschillende
loofbomen: meestal in eiken {Quercus) en beuken {Fagus), maar ook in elzen
{Alnus) en kamperfoelie {Lonicerd) (Du Chatenet 2000: 258-259). A.
cyanescens is vrij gewoon in Europa. Ook in België is dit prachtkevertje niet
zeldzaam.

Phegea 37 (2) (01.VI.2009): 75

Fig. 1. Verspreiding van Agrilus cyanescens Ratzeburg in België (bron K.B.I.N., Brussel).

Agrilus sulcicollis Boisduval & Lacordaire, 1835 (fig. 2)

Tijdens een prospectie te Dilbeek ving ik op 19 mei 2007 met het klopscherm
een exemplaar van Agrilus sulcicollis. Dit staalblauwe prachtkevertje van 7,5
mm viel uit een 40-jarige zomereik ( Quercus robur). De lengte van A. sulcicollis
varieert van 5 tot 8,5 mm. Kenmerkend voor deze Agrilus-soort is het brede
halsschild dat naar de basis toe smaller wordt (Farrugia 2007: 61). De larven
ontwikkelen zich in de schors van eiken ( Quercus ) en beuken ( Fagus ) (Keer
1930: 742). In Midden-Europa is dit prachtkevertje algemeen te vinden van mei
tot augustus (Harde 1979: 235).

Fig. 2. Verspreiding van Agrilus sulcicollis Ratzeburg in België (bron K.B.I.N., Brussel).

Phegea 37 (2) (01.VI.2009): 76

Fig. 3. Verspreiding van Agrilus biguttatus Ratzeburg in België (bron K.B.I.N., Brussel).

Agrilus biguttatus (Fabricius, 1776) (Eikensmalbuikje) (fig. 3)

Op 4 mei 2007 vond ik in de Dilbeekse Wolfsputten achter loszittende schors
van een zomereik ( Quercus robur) een staalblauw kevertje dat zo vast zat in de
schors als een spijker in een plank. Slechts met een zakmes kon het kevertje
losgepeuterd worden. Het bewoog niet en bleek nog in een soort diapauze te
verkeren. Onderzoek van de stukken schors leverde nog 5 andere exemplaren
op. Zij bewogen evenmin. Pas een uur later begonnen de diertjes in het
vangpotje wat rond te kruipen.

De kevertjes werden gemakkelijk gedetermineerd als eikensmalbuikjes,
Agrilus biguttatus. Ze waren alle zes staalblauw van kleur. Toch kan men ook
blauwgroene, groene, messinggroene of olijfgroene exemplaren aantreffen (Keer
1930: 740). Hun lengte bedraagt 9 a 13 mm. Het halsschild is dwarsgerimpeld.
Het schildje (scutellum) vertoont in het midden een scherpingedrukte dwarslijn.
De dekschilden zijn aan het uiteinde afzonderlijk afgerond en zeer fijn getand.
Opvallend zijn de twee viltachtige, witte vlekken in de achterste dekschildhelft.

Op 24 mei 2007 ontdekte ik op dezelfde zomereik nog een exemplaar, maar
ik zag ook twee exemplaren uit het niets komen aanvliegen en neerstrijken op de
stam. Het eerste kevertje was zeer alert. Toen ik naderde, vloog het op en
verdween. Het tweede stoorde zich niet aan mijn aanwezigheid. Het inspecteerde
de schors en begon toen in barstjes en spleetjes telkens één ei af te zetten. Ook
op 4 juni 2007 zag ik op dezelfde stam nog een exemplaar neerstrijken. Later in
het seizoen werden de kevertjes niet meer waargenomen.

Volgens Du Chatenet (2000: 257) ontwikkelen de larven zich niet alleen in
eiken ( Quercus ) maar ook in beuken ( Fagus ) en kastanjebomen ( Castanea ). De
imago's zijn actief in mei en juni. Ze vliegen in de zonneschijn en zijn te vinden
op oude eiken, stronken en geveld hout. A. biguttatus is in Europa nergens
zeldzaam. In België komt hij voor in alle geschikte biotopen.

Phegea 37 (2) (01.VI.2009): 77

Fig. 4. Buprestis novemmaculata Linnaeus, Anderlecht, 25.VIII.1956, leg. E. Derenne, in coll.
K.B.I.N., Brussel.

Buprestis novemmaculata Linnaeus, 1758 (fig. 4)

In het Departement Entomologie van het K.B.I.N. te Brussel bevindt zich een
mooi, afwijkend exemplaar van Buprestis novemmaculata. Het is een robuust
insect van 13 mm lengte. Het etiket vermeldt: "Anderlecht, 25.VIII.1956, leg. E.
Derenne". Deze prachtkever hoort eigenlijk thuis in Oost- en Zuidwest-Europa.
Volgens prachtkeverspecialist M. Neuray gaat het hier zonder twijfel om een
adventief exemplaar dat vermoedelijk in België terecht is gekomen met een
lading naaldhout.

Dankwoord

Informatie en gegevens voor dit artikel werden mij vriendelijk bezorgd door
Remi Guinez (Vorst, Brussel) en Marcel Neuray (Embourg). Voor de
verspreidingskaartjes werd vooral gebruik gemaakt van gegevens uit de
collecties van het K.B.I.N. te Brussel. Hiervoor kreeg ik hulp van Wouter
Dekoninck en Alain Drumont van het Departement Entomologie. Hartelijk dank!

Bibliografie

Du Chatenet, G. 2000. Coléoptères phytophages d'Europe. — N.E.P. Editions, Vitry-sur-Seine.

Harde, K. W. 1979. Buprestidae (Prachtkafer). - In: Freude, Harde & Löhse (eds.) Die Kafer
Mitteleuropas . Band 6. — Goecke & Evers Verlag, Krefeld.

Farragia, S. 2007. Les Agrilus de France. Une clef de détermination. — Magellanes, Andrésy.

Keer, P. M. 1930. Calwer Keverboek. Band 1. — Thieme & Cie, Zutphen.

Troukens, W. 2005. Prachtkevers aan de westrand van Brussel (Coleoptera: Buprestidae). — Phegea
33(2): 76-79.

Troukens, W. 2008. Het beukensmalbuikje, Agrilus viridis (Coleoptera: Buprestidae) aan de westrand
van Brussel. — Phegea 36(3): 101-102.

Phegea 37 (2) (01.VI.2009): 78

Geomyza hackmani (Diptera: Opomyzidae) new for
the fauna of Belgium

Jan Willem A. van Zuijlen

Samenvatting. Geomyza hackmani (Diptera: Opomyzidae) nieuw voor de Belgische fauna.
Gedurende het bestuderen van 56 Belgische Opomyzidae werd een nog niet eerder uit België
gemelde soort aangetroffen: Geomyza hackmani Nartshik, 1984. Het totaal uit België bekende
soorten Opomyzidae komt hiermee op 10.

Résumé. Geomyza hackmani (Diptera: Opomyzidae) une nouvelle espèce pour la faune
beige.

En étudiant 52 exemplaires beiges d’Opomyzidae une espèce a été trouvée qui n’était pas
mentionnée précédemment de la faune beige: Geomyza hackmani Nartshuk, 1984. Ainsi, le
nombre d’espèces d’Opomyzidae beiges s’élève maintentant a 10.

Key words: Geomyza hackmani — faunistics - checklist - Belgium.

Zuijlen, J.W.A. van: Mendelssohnstraat 62, NL-5144 GH Waalwijk, The Netherlands,
JeWe.vZ@Lycos.nl

Opomyzidae is a small acalyptrate family with a mainly Holarctic
distribution. It comprises small (2. 0-4. 5 mm) and rather slender flies with a
yellowish, reddish brown or black body, and long and narrow wings with more
or less distinct maculation. Adults are found in cereal fields, meadows, pastures
and hedges. Larvae develop in the sterns of Poaceae and may cause damage
called the ‘dead heart’. A few species are of economie importance in agriculture
and some of them have numerous host plants.

As in many other countries, this family has received little attention in
Belgium. In 1991 De Bruyn listed seven species of Opomyzidae for Belgium. In
1998 van Zuijlen added three species to this list ( Geomyza martineki Drake,
1992, G. nartshukae Carles-Tolra, 1993, and G. venusta (Meigen, 1830) and
adviced to remove one species (i.e. Geomyza combinata (Linnaeus, 1767)). The
species added by van Zuijlen (1999) were found after studying ca. 950 Belgian
opomyzid specimens deposited in the collection of the Koninklijk Belgisch
Instituut voor Natuurwetenschappen (KBIN). At that time the females of the so-
called combinata group could not be identified reliably and were put aside. In
the meantime it was discovered that the females can be easily identified after the
shape of the 8th abdominal stemite (van Zuijlen 1999).

The female specimens of the combinata group put aside in 1998 have now
been identified: 52 females of G. balachowskyi Mesnil, 1934 - Antwerpen:
Arendonk, Postel, Westmeerbeek; Brabant: Oudergem, Schaffen, Tervuren,
Vilvoorde; Liège: Fringshaus, Hockai, Losheimergraben, Malmédy, Rivage;
Luxembourg: Chiny, Forrières; Namur: Mozet, Rhisnes; Oost-Vlaanderen:
Moorsel, Overmeire; West-Vlaanderen: Knokke; three females of G. hackmani
Nartshuk, 1984 - Moorsel and Ixelles; one female of G. martineki - Pepinster.
G. hackmani has not been recorded from Belgium before. This results in the
following list of species known from Belgium:

Phegea 37 (2) (0 1 . VI.2009): 79

Geomyza Fallén, 1810

balachowskyi Mesnil, 1934
hackmani Nartshuk, 1984
martineki Drake, 1992
nartshukae Carles-Tolra, 1993
tripunctata Fallén, 1823
venusta (Meigen, 1830)

Opomyza Fallén, 1820
florum (Fabricius, 1794)
germinationis (Linnaeus, 1758)
lineatopunctata von Roser, 1 840
petrei Mesnil, 1934

To summarize, G. hackmani is added to the faunal list of Belgium, bringing
the number of Opomyzidae known from Belgium to 10.

I wish to express my gratitude to Dr. P. Grootaert (KB IN) for allowing me to
study the Belgian Opomyzid material.

References

De Bruyn, L. 1991. Opomyzidae. - In: (P. Grootaert, L, De Bruyn & M. De Meyer eds) Catalogue of
the Diptera of Belgium. — Sudiedocumenten van het K.B.I.N. 70: 148.

Zuijlen, J. W. A. van 1998. Three species of Opomyzidae (Diptera: Brachycera) new for the fauna of
Belgium. — Phegea 26(1): 29-30.

Zuijlen, J. W. A., van 1999. Notes on the Fallén collection of Opomyzidae (Diptera) in the
Naturhistoriska Riksmuseet, Stockholm. — Studia dipterologica 6(1): 129-134.

Inhoud:

Asselbergs, J.: Descriptions of new Pyraloidea from the Palaearctic Region (Lepidoptera,

Pyraloidea, Pyralidae: Pyralinae, Phycitinae, Crambinae) 55

Gielis, C.: Nomenclature changes in Afrotropical Pterophoridae (Lepidoptera) 74

Henderickx, FF: A new Feaella species (Pseudoscorpiones: Feaellidae) from Kenya 41

Micevski, B., Micevski, N. & Verovnik, R.: New records of the rare Escher's Blue,

Polyommatus escheri, from the Republic of Macedonia 69

Smit, J. T., van Zuijlen, J. W. A. van & Hanssens, B.: Palloptera scutellata (Diptera:

Pallopteridae) new for the fauna of Belgium 48

Troukens, W.: Nieuwe vondsten van prachtkevers aan de westrand van Brussel

(Coleoptera: Buprestidae) 75

Van de Meutter, F.: Eupeodes goeldlini (Diptera: Syrphidae) new to the Belgian fauna. 51
Zuijlen, V. W. A. van: Geomyza hackmani (Diptera: Opomyzidae) new for the fauna of
Belgium 79

verantw. uitg.: W. De Prins, Dorpstraat 401B, B-3061 Leefdaal (Belgium) - Tel: +32-2-305.37.32
Phegea 37 (2) (01.VI.2009): 80